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In a large skillet, cook the beef and onions over medium heat until no longer pink. Add the garlic and cook for 1 minute longer. Drain off any grease.
Stir in the spaghetti sauce, diced tomatoes, mushrooms and Italian seasoning. Bring the mixture to a boil. Reduce heat and simmer, uncovered, 20 minutes.
Meanwhile, preheat the oven to 350°F. Cook the pasta according to package directions; drain. Editor's Tip: Because the pasta will continue to cook in the oven, you may want to slightly undercook the pasta. Consider cooking for 1 to 2 minutes less than the low end of the pasta's suggested cooking time.
Add the cooked pasta to the beef mixture and gently stir in the sliced plum tomatoes.
Transfer to an ungreased 13x9-in. baking dish. Sprinkle with cheeses. Bake for 25 to 30 minutes or until bubbly and heated through.